I imagine you are loosing a lot of packets trying to communicate via 3 floors on wifi. This means that the destination server is going to keep requesting the missing information until it gets it all. When it is trying to do this during peak usage times it is likely compounding the problem. You said it is not possible to run a line all the way down there but even if you could get a second router on the middle level that might help out a lot. would it be too ugly to run a cable on the outside of your house like on the siding? Just some thoughts.
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